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In 2015, a review recovered mDNA from ancient canid specimens which were discovered on Zhokhov Island plus the Yana river, arctic Siberia. These specimens incorporated the mandible of a 360,000–400,000 YBP Canis cf. variabilis (in which cf. can be a Latin phrase this means unsure). Phylogenetic analyses of such canids discovered 9 mDNA haplotypes

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